Animal shelters in Spain and Belgium have received six lions evacuated from Ukraine, one of which was rescued from a nightclub, the shelters confirmed on Thursday as war rages in the country

The lions, along with six tigers, two wild cats and a wild dog, arrived last week at a zoo in Poland by truck from a refuge near Kyiv following a two-day odyssey skirting battle front lines.

Four of the lions and the wild dog were taken in on Wednesday by a rescue center in Alicante in eastern Spain run by Dutch animal welfare charity AAP, the group said in a statement.

The lions had been living at a shelter near Kyiv after being rescued from "dire" circumstances, with one called Gyz kept in a "small cage in a shopping center."

Another lion called Flori was kept as a pet in a small apartment where it was fed cat food while another named Nila was rescued from a nightclub where it was kept to "entertain customers."

"It will be a challenge to improve their fragile health, step by step, through an adequate diet, exercise and rest," the statement added.

Two other lions named Tsar and Jamil were taken in on Wednesday by the Natuurhulpcentrum shelter in the province of Limburg in eastern Belgium, the group said on Facebook.

The animals will remain in quarantine for three months, it added.

The remaining animals – the six tigers and two wild cats – have so far remained at the zoo in Poznan in western Poland.

The truck, which drove nearly 1,000 kilometers (600 miles) to the Polish border, came face to face with Russian tanks at one point, and had to avoid the Zhytomyr region, which the invading Russian forces have bombarded, a zoo spokesperson said.

At the border, the animals were transferred to a Polish truck while the Ukrainian driver returned home to his children.